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ERX12802066: Single cell RNA sequencing of thymic dendritic cells reveals molecular determinants of antigen transfer in the thymus
4 ILLUMINA (NextSeq 500) runs: 237.9M spots, 21.9G bases, 8.9Gb downloads

Design: Single cell RNA sequencing of thymic dendritic cells reveals molecular determinants of antigen transfer in the thymus
Submitted by: EBI (European Bioinformatics Institute)
Study: Single cell RNA sequencing of thymic dendritic cells reveals molecular determinants of antigen transfer in the thymus
show Abstracthide Abstract
The presentation of self-antigens in the thymus by medullary thymic epithelial cells (mTECs) and dendritic cells (DCs) is crucial for the establishment of central tolerance. While mTECs produce and present self-antigens in an autonomous manner, DCs acquire them from mTECs by Cooperative Antigen Transfer (CAT). Although CAT represents a key process of central tolerance, molecular determinants which drive CAT are largely unknown. To reveal these determinants, we compared the transcriptomes of CAT-experienced and CAT-inexperienced DCs via scRNAseq. Specifically, we sequenced TdTOMATO+ and TdTOMATO– CD11c+ and CD11b+ cells from the thymus of Foxn1CreR26TdTOMATO mice. In this model, fluorescent TdTOMATO protein is exclusively produced by thymic epithelium, thus TdTOMATO expression in DCs (CD11c+ and CD11b+ cells) mark their previous experience of CAT. Using this approach we unraveled several candidate determinants of CAT. We focused on the gene determinant, Cldn1, encoding the tight junction protein Claudin 1, that was found to be highly expressed among the CAT-experienced type 1 lineage DCs. We performed further experiments that confirmed its role in driving CAT. In addition, this scRNAseq approach allowed us to redefine the heterogeneity and maturation states of thymic DCs.
Sample: TdTomato_minus
SAMEA115882145 • ERS20766308 • All experiments • All runs
Organism: Mus musculus
Library:
Name: TdTomato_minus_p
Instrument: NextSeq 500
Strategy: RNA-Seq
Source: TRANSCRIPTOMIC SINGLE CELL
Selection: Oligo-dT
Layout: PAIRED
Construction protocol: Mice were euthanized by cervical dislocation. Thymi were excavated using forceps, cut into 10-15 pieces and digested by enzymatic cocktail of 0.1 mg/ml Collagenase D (Roche) and DNAse I (40 U/ml; Roche) dissolved in RPMI medium supplemented with 3% FBS. Note that pieces of each thymus were put into 1 ml of enzymatic cocktail in 1.5 ml eppendorf tube. To isolate thymic epithelial cells, 0.1 mg/ml Dispase II (Gibco) was added to the enzymatic cocktail. To complete digestion, enzymatic cocktails with thymi were put into thermoshaker, which was set to 37 oC and 800 rpm, and incubated for ~80 minutes. After the incubation, remaining non-digested thymic pieces were pipetted up and down several times using cut tip and the whole solution was transferred into 15 ml falcon tube through filter. Each thymic solution was washed with 2 ml of ice cold 3% FBS and 2 mM EDTA solution in PBS to stop the enzymatic reaction. Then, thymi were spun down (4°C, 400 x g, 10 min). mRNA was extracted from cell suspension during library preparation routine using Chromium next gem single-cell 3′ reagent kit (version 3.1). Cells were resuspended with cocktail of anti-CD11C and anti-CD11B antibodies both conjugated with biotin, stained on ice for 25 min, washed with 3 % FBS and 2 mM EDTA solution in PBS and then spun down (4°C, 300 x g, 10 min). Next, cells were stained with anti-biotin magnetic beads (Miltenyi) according to manufacturer´s protocol and CD11C+ and CD11B+ cells were MACS-enriched using QuadroMACS (Miltenyi). Enriched cells were stained with fluorochrome-conjugated streptavidin on ice for 15 min, washed with 3 % FBS and 2 mM EDTA solution in PBS and then spun down (4°C, 300 x g, 10 min). Next, ~100000 of Streptavidin+ cells were sorted using cell sorter BD Influx (BD Biosciences) from a pool of three littermate thymi. Importantly, half of the cells was sorted as TdTOMATO+ (CAT-experienced cells) and the other half as TdTOMATO– (CAT inexperienced cells) into separate collection tubes to prepare two distinct scRNAseq libraries (samples). To check that viability of sorted cells is higher than 90%, an automated TC20 cell counter (Bio-Rad) was used. ScRNAseq libraries were prepared using Chromium controller instrument and Chromium Next Gen single-cell 3' reagent kit version 3.1 (both 10X Genomics) according to the manufacturer´s protocol targeting 4000 cells per sample. Thus, note that we sequenced 4000 of TdTOMATO+ as well as TdTOMATO– cells.
Runs: 4 runs, 237.9M spots, 21.9G bases, 8.9Gb
Run# of Spots# of BasesSizePublished
ERR1343178360,314,5415.5G2.3Gb2024-08-01
ERR1343178659,297,5195.5G2.2Gb2024-08-01
ERR1343178259,753,5625.5G2.2Gb2024-08-01
ERR1343178758,508,3445.4G2.2Gb2024-08-01

ID:
34518503

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